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The film revolves around a team of trained secret agent animals: guinea pigs Darwin (team leader) (Sam Rockwell), Juarez (martial arts) (Penélope Cruz), Blaster (weapons/transportation) (Tracy Morgan), mole Speckles (cyber intelligence) (Nicolas Cage), and fly Mooch (high-tech surveillance) (Edwin Louis). The team takes on a mission for the US government to stop evil billionaire Leonard Saber (Bill Nighy), who plans to destroy the world with household appliances. Before they can accomplish their mission, the government shuts them down and they are sentenced to a pet shop. Darwin, Juarez, Blaster, Speckles, Mooch, and their new friends Hurley (Jon Favreau) and hamster Bucky (Steve Buscemi) will escape the cage to defeat the villain, stop his threat, and save the world